Australopithecus boisei was first described from a cranium recovered i
n 1959 from Olduvai Gorge, Tanzania(1,2). This and subsequent finds, m
ostly from Kenya's Turkana basin(3-5), resulted in its characterizatio
n as a specialized Australopithecus species with a hyper-robust mastic
atory apparatus(2,4,6). A distinct A. boisei facial morphology has bee
n emphasized to differentiate robust Australopithecus lineages from Ea
st and South Africa(6). A preference for closed and/or wet habitats ha
s been hypothesizes. Here we report some new A. boisei specimens, incl
uding the taxon's first cranium and associated mandible, from Konso, E
thiopia These fossils extend the known geographical range of A. boisei
. They provide clear evidence for the coexistence of A. boisei and Hom
o erectus within a predominantly dry grassland environment. The A. boi
sei specimens from Konso demonstrate considerable morphological variat
ion within the species. The unexpected combination of cranial and faci
al features of this skull cautions against the excessive taxonomic spl
itting of early hominids based on morphological detail documented in s
mall and/or geographically restricted samples.