L. Ginsburg et al., DISCOVERY OF AN ANCESTOR OF THE LESSER PANDA IN THE MIDDLE MIOCENE OFMADRID (SPAIN), Comptes rendus de l'Academie des sciences. Serie II. Sciences de la terre et des planetes, 325(6), 1997, pp. 447-451
One isolated lower tooth (M-2) collected in the Middle Miocene of Madr
id (Spain) is determined as a new taxon: Magerictis imperialensis nov.
gen. nov. sp. It is very close to the lesser panda, Ailurus fulgens.
Thus, the origins of the lesser panda are not with procyonids, but wit
h european mustelids. The origin of ailurids, as well as the simocyoni
ds, are close to a form such as Amphictis a small Oligocene mustelid f
rom the Phosphorites of Quercy, which already shows an elongated M-2.