We consider open supermembranes in an eleven dimensional background. W
e show that, in a flat space-time, the worldvolume action is kappa-sym
metric and has global space-time supersymmetry if space-time has even
dimensional topological defects where the membrane can end. An example
of such topological defects is provided by the space-time with bounda
ries considered by Horava and Witten. In that case the world-volume ac
tion has reparametrisation anomalies whose cancellation requires the i
nclusion of a current algebra on the boundaries of the membrane. The r
ole of kappa-anomalies in a general background is discussed. The tensi
on of the membrane is related to the eleven dimensional gravitational
constant with the aid of the Green-Schwarz mechanism allowing a consis
tency check of M-theory. (C) 1997 Elsevier Science B.V.