Several specimens of Conularia from the mid-Carboniferous Culm Formati
on of the Pyrenees occur as clusters with a large central specimen and
several smaller divergent individuals that may be buds from the large
r central specimen. Other individual and clustered specimens exhibit c
omplex polygonal cross-sections that are inferred to have resulted fro
m longitudinal fission of a precursor pyramid into two or more descend
ant individuals. Apparent budding and transverse fission (strobilation
) of conulariids have been report ed previously, but we have found no
earlier record of longitudinal fission. The existence of this addition
al mode of asexual increase in conulariids further supports their affi
nities with the scyphozoan Cnidaria.