Pa. Allison et Ce. Brett, IN-SITU BENTHOS AND PALEO-OXYGENATION IN THE MIDDLE CAMBRIAN BURGESS SHALE, BRITISH-COLUMBIA, CANADA, Geology, 23(12), 1995, pp. 1079-1082
The Middle Cambrian Burgess Shale was deposited under variable levels
of oxygenation. During periods of oxygenation, low-diversity shell bed
s were formed and the muds were colonized by infauna. Under these circ
umstances pyrite was restricted to anoxic microenvironments and formed
small (1 mm) discrete aggregated masses. Soft-bodied organisms were p
reserved under low-oxygen conditions typified by evenly dispersed pyri
te, and an absence of trace fossils and shell beds.