The magnetospheric convection electric field contributes to Birkeland
currents. The effects of the field are to polarize the plasma by displ
acing the bounce paths of the ions from those of electrons, to redistr
ibute the pressure so that it is not constant along magnetic field lin
es, and to enhance the pressure gradient by the gradient of the bulk s
peed. Changes in the polarization charge during the convection of the
plasma are neutralized by electrons in the form of field-aligned curre
nts that close through the ionosphere. The pressure drives field-align
ed currents through its gradient in the same manner as in quasi-static
plasma, but with modifications that are important if the bulk speed i
s of the order of the ion thermal speed; the variations in the pressur
e along field lines are maintained by a weak parallel potential drop.
These effects are described in terms of the field-aligned currents in
steady state, isotropic, MHD plasma. Solutions are developed by taking
the MHD limit of two-fluid solutions and illustrated in the special c
ase of Maxwellian plasma for which the temperature is constant along m
agnetic field lines. The expression for the Birkeland current density
is a generalization of Vasyliunas' expression for the field-aligned cu
rrent density in quasi-static plasma and provides a unifying expressio
n when both pressure gradients and ion inertia operate simultaneously
as sources of field-aligned currents. It contains a full account of di
fferent aspects of the ion flow (parallel and perpendicular velocity a
nd vorticity) that contribute to the currents. Contributions of ion in
ertia to field-aligned currents will occur in regions of strong veloci
ty shear, electric field reversal, or large gradients in the parallel
velocity or number density, and may be important in the low-latitude b
oundary layer, plasma sheet boundary layer, and the inner edge region
of the plasma sheet.