Pollen, spores, and other plant microfossils are important constituent
s of Tertiary coals, both as petrographic components (the maceral spor
inite) and as distinctive indicators of the nature of the depositional
environment in which the coal-forming Feat accumulated. Palynological
analyses, which have been conducted on Tertiary coals since the 1930'
s, play an increasingly important role in modem paleoecological analys
es of coals. Among the most important factors controlling the petrogra
phic composition and facies characteristics of coal deposits are the t
ypes and relative abundances of plants that composed the peat. At pres
ent, wetland environments in which peat accumulates (mires) range from
marshes and bogs to forested swamps; each is inhabited by distinctive
plant communities. These plant communities are associated with varied
conditions of climate, water depth, chemistry, and nutrient supply, a
nd the floras can be indicative of these conditions. In the Tertiary P
eriod, plant communities of mires were also highly diverse and special
ized (especially as compared with those of the Carboniferous). The dom
inant species in some Paleocene mire floras were gymnosperms, but cont
inuing a trend begun in Late Cretaceous time, angiosperms became incre
asingly prominent through the Tertiary. The diverse angiosperm-dominat
ed mire floras produced coals that vary significantly in facies and he
nce in quality. Such contrasts are evident in contemporaneous coals fr
om different paleolatitudes, as well as among coals of different ages
within the Tertiary, as exemplified by selected deposits in the wester
n United States. The palynofloras of these coals reflect the floristic
composition of the plant communities of the ancient mires.