Jm. Adrain et Bde. Chatterton, THE OTARIONINE TRILOBITE CYPHASPIS, WITH NEW SPECIES FROM THE SILURIAN OF NORTHWESTERN CANADA, Journal of paleontology, 70(1), 1996, pp. 98-108
The genus Cyphaspis Burmeister, 1843, is first known from the northern
Laurentian Sheinwoodian. By the late Homerian, the genus had appeared
in England and Baltica, and from the Ludlow through the Lower and Mid
dle Devonian it had an essentially cosmopolitan distribution. The Miss
issippian Dixiphopyge Brezinski, 1988, may represent a relict distribu
tion of Cyphaspis, and if so should be considered a junior subjective
synonym. Cyphaspis is considered the sister taxon of Otarion Zenker, 1
833. The oldest, Sheinwoodian, species of either genus are very simila
r, but the clades evolved to the point of gross morphological disparit
y in the Devonian. New species from the Wenlock and probably Ludlow of
northwestern Canada are Cyphaspis lowei, C. munii, C. buchbergeri, an
d C. mactavishi. New ontogenetic material of the Zlichovian C. dabrown
i (Chatterton, 1971) further demonstrates the pervasiveness of the bas
ic juvenile morphology of the tribe Otarionini.