Many constructional and taphonomic incongruencies dissolve if ammonite
s are modeled as active Cartesian divers, whose last septum remained a
rubber-like, ulcalcified membrane until the next chamber cycle began.
If the prochoanitic invagination of the septal mantle served as a gas
gland able to produce gas at or above ambient pressure, the chamber b
ehind could be operated like a swim bladder by sutural muscles spannin
g across the septal mantle. Other septal functions probably served as
evolutionary stepping stones on the way to this unique design.