J. Adachi et al., PHYLOGENETIC-RELATIONSHIPS OF THE BERBERIDACEAE BASED ON PARTIAL SEQUENCES OF THE GAPA GENE, Plant systematics and evolution, 1995, pp. 351-353
A Phylogenetic tree of six species in the Berberidaceae based on parti
al sequences of the nuclear gene for glyceraldehyde-3-phosphate dehydr
ogenase (gapA) shows that the Berberidaceae is a distinctly monophylet
ic group, and divided into three clades:Nandina-Caulophyllum, Mahonia-
Ranzania, Epimedium-Diphylleia.