Because a large number of theoretical models suggest chaos in populati
ons, held biologists have been trying for decades to confirm the exist
ence of chaos in nature. In spite of their efforts, chaotically evolvi
ng populations have been found in extremely low numbers. In this artic
le we consider a metapopulation model which was built up by the intera
ction of local populations. Local populations interact with their near
est neighbours via migrations, but migration occurs only if the local
population density exceeds a threshold level (overcrowding). Depending
on the strength of the interaction, the metapopulation density shows
noiselike dynamics of many degrees of freedom, periodical evolution, o
r tends to a fixed point. Low dimensional collective chaos has not bee
n detected. Moreover, the migration size distribution indicates the em
ergence of self-organized criticality, if the interaction is strong en
ough.