Wp. Coombs et Ta. Demere, A LATE CRETACEOUS NODOSAURID ANKYLOSAUR (DINOSAURIA, ORNITHISCHIA) FROM MARINE-SEDIMENTS OF COASTAL CALIFORNIA, Journal of paleontology, 70(2), 1996, pp. 311-326
A partial nodosaurid ankylosaur skeleton, consisting primarily of the
ilia, hindlimbs, posterior dorsal armor, plus partial forelimb element
s and additional armor, was recovered from the marine Point Loma Forma
tion, late Campanian age, north of San Diego, California. The specimen
is similar to contemporaneous species of Panoplosaurus and Edmontonia
from terrestrial sediments of the western interior, but there are als
o similarities to the armor of Stegopelta landerensis from marine sedi
ments of earliest Cenomanian age from Wyoming. Skeletal elements criti
cal for generic determination are not preserved, and the specimen is i
dentified as Nodosauridae, incertae sedis. An associated marine invert
ebrate fauna and nannoplankton flora have revealed some discrepencies
in the correlation systems used for Upper Cretaceous marine rocks of c
oastal California. Hollow limb bones of the specimen are interpreted a
s a preservational artifact. Nodosaurids had broad ecological toleranc
es, and visited riparian and coastal environments more frequently than
other dinosaurs. However, a review of morphologic and distributional
evidence fails to support a theory of amphibious or aquatic habits for
nodosaurids.