I studied ecology and behavior of wintering Black-throated Blue Warble
rs (Dendroica caerulescens) at three sites on Puerto Rico over four ye
ars. The site with tall mature forest and relatively few fruiting unde
rstory plants had warbler populations characterized by: a predominance
of males; relatively early fall arrival of returning adults; high sit
e fidelity; large home ranges; low density of sedentary birds; few wan
dering individuals; high over-winter persistence of sedentary birds; a
nd an invertebrate-rich diet. In contrast, the shrubby second-growth s
ite with an abundance of fruiting plants had warbler populations chara
cterized by: a predominance of females; relatively late arrival of ret
urning adults compared to juveniles; low site fidelity; small home ran
ges; high density of sedentary and wandering birds; low overwinter per
sistence of sedentary birds; and a nectar- and fruit-rich diet. At the
third site, the population trails fell within these extremes. Some si
te variation is attributable to sex differences in site persistence, w
andering, and home-range size, which appear as population differences
only because sex ratios vary among sites. However, differences in abun
dance and seasonality of fruit and nectar may further contribute to si
te differences in overwinter persistence of sedentary birds, wandering
, home-range size, and density.