Lj. Musselman et al., ISOETES X BRUNTONII (ISOETES-ENGELMANNII X ISOETES-HYEMALIS), A NEW HYBRID QUILLWORT FROM VIRGINIA, American fern journal, 86(1), 1996, pp. 8-15
Isoetes xbruntonii is the hybrid between the diploid I. engelmannii (2
n=22) and the tetraploid I. hyemalis (2n=44). An apparently sterile tr
iploid (2n=33), it has megaspore features intermediate between the two
parents, bur does not exhibit hybrid vigor. Isoetes xbruntonii is fre
quent at localities in the outer Coastal Plain of Virginia.