The acoustic echo levels from zooplankton are strongly dependent upon
the acoustic frequency and size, shape, orientation, and material prop
erties of the animals. Because of the great number of species of zoopl
ankton, it is practical to study the acoustic properties of species gr
ouped by their gross anatomical similarity. Zooplankton from several m
ajor groups are discussed: fluid-like (decapod shrimp, euphausiid, sal
p), hard elastic shelled (gastropod), and gas-bearing (siphonophore).
The results from laboratory tests show that the plots of (single ping)
target strength versus acoustic frequency have a distinct pattern for
each animal type. For example, the plot for euphausiids when ensonifi
ed at broadside incidence contained a series of broadly spaced deep nu
lls; the plot for gastropods either had more tightly spaced nulls or a
flat spectrum; the plot for siphonophores either had a less consisten
t pattern of nulls or a flat spectrum. The nulls from the euphausiid d
ata were sometimes as deep as 30 dB below surrounding levels. The patt
erns are linked to the physics of the scattering process and modeled m
athematically. In addition, key results on these animals from Stanton
et al. (1994a, ICES J. Mar. Sci., 51: 505-512) are summarized to furth
er illustrate the variability in scattering characteristics of the ani
mal groups (for example, data from 2-mm-long gastropods show that they
produce a level of echo energy per unit biomass approximately 19 000
(i.e., 43 dB) times greater than that of 30-mm-long salps). The impact
of these observations on design and interpretation of acoustic survey
s is discussed. Very importantly, drawing a simple relationship betwee
n echo energy and biomass for regions containing a complex assemblage
of zooplankton would be greatly flawed. (C) 1996 International Council
for the Exploration of the Sea