Black Pine (Pinus nigra Arnold) is naturally distributed on xerotherm
limestone and silicate sites of the mediterranean and submediterranean
regions and is used for artificial afforestations of xerotherm waste
land, dry pastures and dunes in more northern regions. Waste land and
dry pastures are frequently found on limestone sites. This would sugge
st that Black Pine requires limestone sites. However, nutrition experi
ments have shown that this is nor the case. Black Pine has a low need
of nutrients; there was no lack of nutrients on any investigated site
in Germany, the Czech Republic, Slovakia, Hungary and Georgia (Caucasu
s). On the other hand, water supply was found to be the growth limitin
g factor. The element concentrations in the needles of Black Pine are
mostly lower than those of Scotch Pine (Pinus sylvestris L.) under the
same conditions, except for Potassium concentration, which is higher
in Black Pine needles. In pot experiments with sufficient water supply
nutrition and growth can be improved by fertilisation. On natural, of
ten extremely dry sites typical of Black Pine stands fertilisation has
no effect on growth rates, due to minimum water supply.