The general characteristics of global vegetation during the middle Pli
ocene warm period can be reconstructed from fossil pollen and plant me
gafossil data. The largest differences between Pliocene vegetation and
that of today occurred at high latitudes in both hemispheres, where w
arming was pronounced relative to today. In the Northern Hemisphere co
niferous forests lived in the modern tundra and polar desert regions,
whereas in the Southern Hemisphere southern beech apparently grew in c
oastal areas of Antarctica. Pliocene middle latitude vegetation differ
ed less, although moister-than-modem conditions supported forest and w
oodland growth in some regions now covered by steppe or grassland. Pli
ocene tropical vegetation reflects essentially modern conditions in so
me regions and slightly cooler-than-or warmer-than-modern climates in
other areas. Changes in topography induced by tectonics may be respons
ible for many of the climatic changes since the Pliocene in both middl
e and lower latitudes. However, the overall latitudinal progression of
climatic conditions on land parallels that seen in the reconstruction
of middle Pliocene sea-surface temperatures. Pliocene paleovegetation
al data was employed to construct a 2 degrees x 2 degrees global grid
of estimated mid-Pliocene vegetational cover for use as boundary condi
tions for numerical General Circulation Model simulations of middle Pl
iocene climates. Continental outlines and topography were first modifi
ed to represent the Pliocene landscape on the 2 degrees x 2 degrees gr
id. A modem 1 degrees x 1 degrees vegetation grid was simplified and m
apped on this Pliocene grid, and then modified following general geogr
aphic trends evident in the Pliocene paleovegetation data set.