M. Kottelat, THE IDENTITY OF PUNTIUS-EUGRAMMUS AND DIAGNOSES OF 2 NEW SPECIES OF STRIPED BARBS (TELEOSTEI, CYPRINIDAE) FROM SOUTHEAST-ASIA, The Raffles bulletin of zoology, 44(1), 1996, pp. 301-316
The synonymy of Puntius eugrammus and P. johorensis is confirmed. Four
species of barbs of the genus Puntius with longitudinal stripes occur
in Southeast Asia: P. lineatus with 0 or 2 barbels and fleshy lips wi
th a continuous postlabial groove, from the Malay Peninsula, Sumatra a
nd western Borneo; P. johorensis with four barbels and a juvenile colo
ur pattern of 4 vertical bars changing into 5-6 stripes at about 20-30
mm SL, from the Malay Peninsula, Sumatra, and Borneo; P. trifasciatus
, new species, with four barbels and a juvenile colour pattern of 4 ve
rtical bars changing into 3(-4) stripes at about 30-40 mm SL, from wes
tern and southern Borneo; and P. gemellus, new species, with four barb
els and juveniles with 4-5 irregular vertical bars changing into 5-6 n
arrow stripes at about 18-20 mm SL, from southern Sumatra, Bangka and
southern Borneo.