Sg. Lucas et al., MARINE FOSSIL SHARK (CHONDRICHTHYES) FROM NONMARINE EOCENE SEDIMENTS,NORTHEASTERN KAZAKHSTAN, Proceedings of the Biological Society of Washington, 109(2), 1996, pp. 349-352
A single tooth of the sand tiger shark, Carcharias sp., was collected
from the upper Eocene Kusto svita in the Zaysan basin of northeastern
Kazakhstan. The Kusto svita is nonmarine lacustrine strata with an ext
ensive nonmarine biota of plants (especially charophytes) and terrestr
ial vertebrates (especially mammals). The shark tooth was apparently c
arried to the site by a predator/scavenger from the then-nearest seawa
y, more than 500 km to the northwest.