Gw. Storrs et al., THE DIAPSID REPTILE, PACHYSTROPHEUS-RHAETICUS, A PROBABLE CHORISTODERE FROM THE RHAETIAN OF EUROPE, Palaeontology, 39, 1996, pp. 323-349
The enigmatic Upper Triassic reptile, Pachystropheus rhaeticus, displa
ys characters suggestive of choristoderan affinity and, as such, is po
tentially the oldest known choristoderan reptile. Examination of the k
nown skeletal elements indicates that the choristoderan lineage remain
ed morphologically conservative throughout its recorded history. The o
ccurrence of Pachystropheus fossils in marginal marine bone beds, howe
ver, may reflect a previously unrecognized shift of habitat for the Ch
oristodera, from the paralic environments occupied by early representa
tives, to the more typical freshwater, often fluvial, deposits contain
ing later forms.