Feathers are subject to large aerodynamic forces during fight and must
therefore be stiff and strong, yet also light. Published estimates of
Young's modulus of the feather keratin composite vary widely in the l
iterature, and it is suggested that this is predominantly due to test
methodology rather than true interspecific differences. The factors in
fluencing the Young's modulus of keratin are discussed, together with
their functional implications. Damage resistance must also be good; fe
athers that are lost or broken can only be replaced infrequently, but
regularly, at moult. Feather shafts appear to be very strong for their
weight. The possible role of the medullary foam in preventing bucklin
g failure is still unclear; medulla removal only appears to have a min
imal effect on both the stiffness and strength of whole sections of th
e rachis. Melanic keratin has properties that make it more competent a
t resisting abrasive wear than non-melanic keratin.