Local adaptation of insect herbivores to individual variation of host
plant characteristics (i.e. deme formation) may be counteracted by gen
e flow among those hosts. The winter moth lives on oak trees which bud
burst dates vary considerably among individuals. Winter moth egg hatch
shows a synchronization with individual oak budburst, which is partia
lly promoted by the female's brachyptery and thus low dispersal. Males
are, however, winged and capable of flying to other trees. The mark-r
elocation experiments reported in this paper show that males also exhi
bit very limited dispersal, possibly increasing egg hatch synchronizat
ion and expected fitness. The potential importance of the wings for ma
les is discussed.