PHYLOGENY OF THE INTERTIDAL ALEOCHARINE TRIBE LIPAROCEPHALINI (COLEOPTERA, STAPHYLINIDAE)

The tribe Liparocephalini (genera Liparocephalus Maklin, 1853; Diaulot a Casey, 1893 [ = Genoplectes Sawada, 1955]; Paramblopusa Ahn & Ashe, Amblopusa Casey, 1893; Salinamexus Moore & Legner, 1977 [ = Biophytosu s Moore & Legner, 1977]), all of which are exclusively restricted to t he Pacific coasts of the Holarctic Region, is hypothesized to be a mon ophyletic group based on the following synapomorphies: seta v absent ( inferred to be lost) from mentum (reversed one time in the Liparocepha lus lineage), setae distributed only on mesal surface of galea and ape x with setae, one medial seta present on prementum, and contiguous mes ocoxal cavities. Natural history and history of the classification of the Liparocephalini are discussed. Phylogenetic analyses of the specie s of the Liparocephalini are presented based on larval (21 characters, 57 states), adult (49 characters, 115 states), and a combination of l arval and adult characters (70 characters, 172 states). Analysis of th e combined larval and adult data sets with successive approximation re sulted in a single most parsimonious tree (length = 937, CI = 0.885, R T = 0.930) with the following patterns of generic relationships (outgr oup (Salinamexus + Biophytosus (Amblopusa (Paramblopusa (Diaulota + Ge noplectes, Liparocephalus))))).