E. Diazmartinez et al., FIRST RECORD OF ARTHRODIRE (PLACODERMI, V ERTEBRATA) FROM THE DEVONIAN OF SOUTH-AMERICA, Comptes rendus de l'Academie des sciences. Serie II. Sciences de la terre et des planetes, 323(4), 1996, pp. 349-356
Dermal plates of a large eubrachythoracid arthrodire (Placodermi, Vert
ebrata), provisionally referred to a dunkleosteid, are recorded from t
he detrital facies of the Famennian Colpacucho Formation, in the Cuman
a Peninsula (Lake Titicaca, Bolivia). This is the first occurrence of
arthrodires in the Devonian of South America. These arthrodire plates
are found in association with a large chondrichthyan spine which recal
ls some of the Famennian and Early Carboniferous species of ''Ctenacan
thus''.