WHEN young birds leave on their first migration, they are guided by in
nate information about their direction of migration. It is generally a
ssumed that this direction is represented twice, namely with respect t
o celestial rotation and with respect to the Earth's magnetic field(1,
2). The interactions between the two cue systems have been analysed by
exposing hand-raised young birds during the premigratory period to cu
e-conflict situations, in which celestial rotation and the magnetic fi
eld provided different information. Celestial rotation altered the cou
rse with respect to the magnetic field(3-7), whereas conflicting magne
tic information did not seem to affect the course with respect to the
stars(8,9). Celestial information thus seemed to dominate over magneti
c information. Here we report that the interaction between the two cue
systems is far more complex than this. Celestial rotation alone seems
to provide only a tendency to move away from its centre (towards geog
raphical south), which is then modified by information from the magnet
ic field to establish the distinctive, population-specific migratory d
irection.