More detailed observations of the synaptonemal complex (SC) in asynapt
ic maize plants have been faciliated by superior silver-staining proce
dures. These suggest that central region components of the SC are stro
ngly implicated as defective in asynaptic. Apparently homologous axial
elements tend to follow roughly parallel courses within the nucleus a
t pachytene, in some short segments apparently synapsed and in others
at wider separation than normal synapsis yet close enough to allow obs
ervation of thin central element segments and also occasional thin tra
nsverse element-type structures. This kind of transverse filament may
be weakened and severely stretched yet associated with both axial elem
ents. Small nodules, similar to recombination nodules, appear at corre
sponding positions in widely separated axial elements.