Wr. Branch et al., A REVIEW OF THE NAMAQUA GECKO, PACHYDACTYLUS-NAMAQUENSIS (REPTILIA, GEKKONIDAE) FROM SOUTHERN AFRICA, WITH THE DESCRIPTION OF 2 NEW SPECIES, South African journal of zoology, 31(2), 1996, pp. 53-69
An analysis of morphological and allozyme variation in the Namaqua gec
ko, Pachydactylus namaquensis from southern Africa is presented. Three
separately evolving lineages, well defined by morphology and allozyme
variation, are identified. The isolated southern population, occurrin
g on the southern escarpment and Cape Fold Mountains surrounding the w
estern Little Karoo, is named P. kladaroderma sp. nov., and is charact
erized by a slit-like ear opening, low number of granules bordering th
e mental (3-6) and mental and adjacent infralabials (5-13), the freque
nt (79%) occurence of the supralabial entering the nostril, and its dr
ab brown base colouration. A northern population, occurring in souther
n Namibia and the Richtersveld is named P. haackei sp. nov., and is ch
aracterized by its more rounded or squared ear opening; high number of
granules bordering the mental and adjacent infralabials (11-19), the
general exclusion of the supralabial from the nostril (only 3.7% entry
), and brighter, lighter colouration. It is further differentiated fro
m P. kladaroderma on the basis of fixed differences at 11 allozyme loc
i. Both new species differ from P. namaquensis, which is mainly restri
cted to Little Namaqualand, but is sympatric with P. haackei in the Lo
wer Orange River region, by their more heterogenous dorsal scalation,
smaller cloacal spurs, lack of spine-like tubercles on the lateral sur
faces of the tail, and more fragile skin. The type locality of P. nama
quensis is restricted to 'the vicinity of Springbok, Little Namaqualan
d, Northern Cape Province, South Africa'.