Spatial patterns in species richness of the complete New World avifaun
a were analysed, using data previously employed to examine spatial tre
nds in geographic range size. This allowed variation in the patterns t
o be compared. Species richness was highest around the equator, and de
creased towards higher latitudes in both hemispheres. This decrease wa
s asymmetrical; at equivalent latitudes, richness was higher in the so
uthern than in the northern hemisphere, although the reverse was true
for a measure of endemism. Controlling for latitude, species richness
was higher in the west than in the east. The net primary productivity
of, and solar radiation received by an area were both correlated with
species richness. However, neither explained more variation in richnes
s than did latitude. No single mechanism developed to explain spatial
patterns in species richness satisfactorily explains the patterns obse
rved in the New World avifauna. We discuss reasons why this might be t
he case. Finally, we point out that species richness at low latitudes
is not simply a multiplication of richness at high latitudes; species
found at high and low latitudes are unlikely to be ecologically equiva
lent. Any mechanism that is proposed to explain richness patterns in N
ew World birds will need also to account for this observation.