Further progress in understanding insect cold-hardiness requires a bro
ader perspective than hitherto. Such a broader approach is feasible, g
iven the range of information now available on cold-hardiness and the
variety and quality of equipment to study it. Comprehensive informatio
n on individual species is required, using the full range of available
techniques, instead of piecemeal investigations. Comparisons of speci
es must likewise be based on how their complete cold-hardiness strateg
ies are structured, rather than on any particular contrasts between in
dividual elements. Most importantly, a well based understanding of col
d-hardiness requires wider linkages between cold-hardiness and other s
alient aspects of the life cycle, such as seasonal timing and control,
water balance, and metabolism and energy budgets. Cold-hardiness is o
nly one aspect of the adaptive trade-offs that structure the life cycl
e of any given species.