Environmental crises provide ecology with a new impetus. Indeed, acid
rain, massive changes in land-use, and the prospect of global climate
change are giving the study of ecosystems a new meaning and new mandat
es. Careful natural history descriptions are no longer sufficient for
evaluating potential impacts of changing environmental conditions. Und
erstanding how ecosystems work has become a priority. The present pape
r investigates the role of water stress as an agent of rapid vegetatio
n change. A sensitivity study is performed to explore the issue. Envir
onmental changes are imposed, rather than obtained from atmospheric ge
neral circulation model (GCM) output, to facilitate the interpretation
of the results, to enable the identification of generic patterns, in
particular, critical thresholds, and to foster the understanding of un
derlying processes. A physically- and physiologically-based, climatica
lly-sensitive, numerical simulation model of forest dynamics, the Ener
gy, Water, and Momentum Exchange and Ecological Dynamics (EXE) model,
is used for this purpose. The results of the sensitivity analysis unde
rtaken with EXE document the interplay between physical and physiologi
cal feedbacks and ecological forest responses to climatic changes. Mor
eover, they explain the relative importance of temperature, water, and
nitrogen limitations, indicating that water stress is responsible for
some of the fastest vegetation changes.