Carbon storage and flow through forest ecosystems are major components
of the global carbon cycle. The cycle of carbon is intimately coupled
with the cycle of nitrogen and the flow of water through forests. The
supply of water for tree growth is determined by climate and soil phy
sical properties. The rate at which nitrogen mineralization occurs dep
ends on climate and the type of carbon compounds with which the nitrog
en is associated. Species composition, which is also affected by clima
te, can greatly influence the composition of carbon compounds and subs
equently nitrogen availability. Climate change can therefore have a di
rect effect on forest ecosystem production and carbon storage through
temperature and water limitations, and an indirect effect through the
nitrogen cycle by affecting species composition. Model simulations of
these interactions show that climate change initiates a complex set of
direct and indirect responses that are sensitive to the exact nature
of the project climate changes. We show results using four different c
limate-change projections for a location in northeastern Minnesota. Mo
deled forest responses to each of these climate projections is differe
nt indicating that uncertainties in the climate projections may be amp
lified further as a result of shifts in balance between positive and n
egative ecosystem feedbacks.