Factors influencing the likelihood that a coyote, Canis latrans, will
disperse or remain in its natal pack are not well understood. The soci
al and nutritional factors influencing the dispersal of resident coyot
es in Yellowstone National Park, Wyoming were examined by observing 49
coyotes from five resident packs for 2456 h from January 1991 to June
1993. One of two strategies was adopted: disperse or remain in the na
tal pack (philopatry). Dispersing coyotes were low-ranking pups, or lo
w-ranking betas, who spent little time with other pack members and wer
e typically subordinate during interactions with other members of thei
r pack. Dominant (alpha) coyotes and higher-ranking betas did not disp
erse. Dispersers had little or no access to ungulate carcasses during
winter compared with higher-ranking, philopatric individuals in the pa
ck. The ability to capture small mammals also became important in dete
rmining which animals remained or dispersed. When pack size increased
in the winter of 1992-1993, possibly intensifying competition at ungul
ate carcasses, philopatric pups and betas captured small mammals at a
higher rate than dispersing coyotes. Individuals that remained in the
pack were dominant and higher-ranking, typically had greater access to
carcasses in their respective packs and captured small mammals at a h
igher rate than dispersing individuals when pack sizes were largest. P
hilopatric coyotes remained within their pack with the objective of ad
vancing to the alpha breeding position. Low-ranking, subordinate coyot
es left their natal pack and attempted to establish themselves in eith
er adjacent or distant territories. (C) 1996 The Association for the S
tudy of Animal Behaviour