TIDAL AND SEASONAL-CHANGES IN THE TEMPORAL AND SPATIAL-DISTRIBUTION OF FORAGING CARCINUS-MAENAS IN THE WEAKLY TIDAL LITTORAL-ZONE OF KERTEMINDE FJORD, DENMARK

Seasonal and tidal changes in foraging activity of the shore crab Carc inus maenas were investigated from August 1992 to May 1993 in Kertemin de Fjord, Funen, Denmark. Foraging activity was 15 to 20 times higher in summer and autumn than in winter and spring. Despite the weakly tid al conditions at the study site a significant proportion of the crab p opulation exhibited tidal rhythmicity in foraging activity. Detailed e xamination revealed that it was predominantly early intermoult (green ventral carapace), male crabs that behaved in this way. Females consti tuted ca 20% of the crabs caught throughout the year and were active p rimarily at night, independent of the tidal phase. In constant conditi ons in the laboratory, the majority of male crabs displayed endogenous heart rate rhythms in phase with expected tidal and diel cycles at th eir collection sites. Non-green male crabs did not exhibit rhythmicity to the same degree. Juvenile crabs comprised a greater proportion of the total catch at low tide than al high tide. The findings are compar ed with earlier studies on C. estuarii (earlier C. mediterraneus) in s imilar tidal conditions in the Mediterranean Sea.