TERRESTRIAL arthropods have a fossil record that reaches back to at le
ast the Upper Silurian (420 million years ago). Most available data on
these early land animals comes from a few sites at which abundant mic
roarthropod remains are preserved as organic cuticle fragments. Such c
uticle assemblages imply that early terrestrial ecosystems were domina
ted by small arthropods of the soil and litter, We describe here new i
mpression and impression/compression fossils from the Emsian stage of
New Brunsnick and Quebec, Canada, which include millipeds, arthropleur
ids, and a scorpion with book-lungs preserved. They suggest that surfa
ce macroarthropods were also a significant component of the land fauna
.