Gc. Williams, LIVING GENERA OF SEA PENS (COELENTERATA, OCTOCORALLIA, PENNATULACEA) - ILLUSTRATED KEY AND SYNOPSES, Zoological journal of the Linnean Society, 113(2), 1995, pp. 93-140
An illustrated dichotomous kev and synopses of the 32 genera of living
pennatulacean octocorals are presented, which incorporate new morphol
ogical and distributional data from the examination of recently collec
ted material. In addition, a key to the 15 extant families, lists of v
alid genera, synonyms, and a table of comparative characters are also
included. Lastly, a revised classification and phylogenetic considerat
ions are presented. Preliminary investigations indicate that the tradi
tional higher classification scheme of the Pennatulacea is inadequate
for reasons of paraphyly and intermediate taxa, that tend to negate pr
ecise distinctions between some of the nominal higher taxa. Of the app
roximately 436 described species of sea pens worldwide, only 186 (or 4
3%) are estimated to be valid. In addition, several undescribed specie
s have recently been discovered, and others will no doubt be discovere
d in the future. It is therefore estimated that tile extant pennatulac
ean fauna of the world comprises approximately 200 species in 32 gener
a.