Bryophytes, including the mosses, liverworts, and hornworts, occur in
a variety of habitats with high concentrations of metals and have othe
r characteristics that are advantageous for studies of metal tolerance
. Mosses may evolve genetically specialized, metal-tolerant races less
frequently than flowering plants. Some species of mosses appear to ha
ve inherently high levels of metal tolerance even in individuals that
have not been subjected to natural selection in contaminated environme
nts. Scopelophila cataractae, one of the so-called copper mosses, not
only tolerates extremely high concentrations of metals in its substrat
es, but requires these substrates for optimum growth. This species sho
uld be included in mechanistic studies of tolerance at the cellular an
d molecular levels.