W. Streit et D. Fengel, FORMATION AND DEPOSITION OF TANNINS IN QUEBRACHO-COLORADO (SCHINOPSIS-BALANSAE ENGL), Holz als Roh- und Werkstoff, 53(1), 1995, pp. 56-60
In Quebracho colorado heartwood tannins are located in the lumina of m
ost parenchyma cells, fibres and vessels. Formation and deposition of
the tannins were studied by electron microscopy. In parenchyma cells p
henolic compounds first occur at the inner and outer side of the membr
anes of vacuoles. Later, these vacuoles are completely filled with dar
k stained compounds and fuse together. From parenchyma cells the pheno
lic substances migrate into the lumina of fibres and vessels through p
its. In pits they penetrate into the compound middle lamella (CML) and
diffuse along the CML. Tannins were also found in the cell walls and
in fissures within the CML and G-layer of tension wood fibres.