A nonlinear demographic model was used to predict the population dynam
ics of the flour beetle Tribolium under laboratory conditions and to e
stablish the experimental protocol that would reveal chaotic behavior,
With the adult mortality rate experimentally set high, the dynamics o
f animal abundance changed from equilibrium to quasiperiodic cycles to
chaos as adult-stage recruitment rates were experimentally manipulate
d. These transitions in dynamics corresponded to those predicted by th
e mathematical model. Phase-space graphs of the data together with the
deterministic model attractors provide convincing evidence of transit
ions to chaos.