YIELD RESPONSES OF 2 WHEAT GENOTYPES TO CARBON-DIOXIDE AND TEMPERATURE IN-FIELD STUDIES USING TEMPERATURE-GRADIENT TUNNELS

Clear, plastic-coated, temperature gradient tunnels (TGTs), 8 X 1.25 X 1.25 m were designed and built to examine how temperature and CO2 aff ect the yield of wheat in the field. Each of the three modules of each TGT was maintained at a different temperature above the ambient tempe rature using solar heating during the day and electric heating at nigh t. The maximum day-time increment above ambient for the warmest module was 5 degrees C and full-season averages were close to 2 degrees C. T GTs were paired, with air in one being enriched to 700 mu L L(-1) CO2, and in the other being maintained at ambient CO2. Crops were planted in the TGTs at two sites in either summer (December) or winter (April and July) and they remained there until maturity. CO2 enrichment incre ased the yield in summer plantings by up to 36%. In winter plantings, with mean temperatures between sowing and anthesis of around 10 degree s C, the responses to CO2 were small averaging only 7% (range 1-12%). Though yield declined with increasing temperature in the TGTs in summe r, there was a clear trend for an increasing response to CO2 at these higher temperatures, i.e. yield declined less. In summer, there was no convincing evidence for a different relative response to CO2 in two i solines which differed in maturity date, though the later line yielded more under the highest temperature regime (mean of 22-24 degrees C be tween sowing and anthesis). In winter there was a strong trend for the isoline requiring less vernalisation to respond more to CO2. It is su ggested that early progress towards flowering might predispose wheat t o a greater CO2 response. Overall, the data indicated that the positiv e response to CO2 in grain yield is likely to increase at approximatel y 1.8% per 1 degrees C in wheat crops that are not limited by water. E xtrapolation indicated that the temperature at which there was no resp onse to CO2 was 5 degrees C. All yield responses reflected biomass res ponses as harvest index was unchanged by CO2.