PHYLOGENETIC-RELATIONSHIPS OF THE GENERA OF THE PRONOCEPHALIDAE LOOSS, 1902 (DIGENEA, PARAMPHISTOMIFORMES)

Quantitative phylogenetic analysis of 20 nominal genera of the Pronoce phalidae based on 47 morphological transformation series produced 6 eq ually parsimonious trees, each with a consistency index of 77.8%. All trees agree that Adenogaster is the sister group to the rest of the pr onocephalids, and a new subfamily is proposed for it. The Pronocephali nae comprises Pronocephalus, Ruicephalus, Neopronocephalus, Macravesti bulum, Choanophorus, Cetiosaccus, and Metacetabulum. The Charaxicephal inae comprises Charaxicephalus, Desmogonius, Diaschistorchis, Pleurogo nius, Iguanacola, Renigonius, Parapleurogonius, Himasomum, Pyelosomum, Cricocephalus, Barisomum, and Pseudobarisomum. An amended diagnosis f or Himasomum is presented. The trees differ only in the placements of Pleurogonius, Renigonius + Parapleurogonius, Iguanacola, and Himasomum relative to each other. Parapronocephalum and Notocotyloides are memb ers of the clade containing the Notocotylidae. The phylogenetic tree s upports interpretations of 3-4 transitions from marine to freshwater t urtles, 3 host switches from marine turtles to the Galapagos marine ig uana and 3 from marine turtles to the French angelfish, and widespread host switching among marine chelonians. No switches to non-chelonian hosts coincide with transitions from marine to freshwater.