AN exceptionally preserved new specimen of the giant conodont Promissu
m pulchrum reveals details of the trunk musculature, feeding apparatus
and eyes, High-fidelity resolution of ultrastructural features of the
trunk myomeres provides the first conclusive evidence of muscle-fibre
organization and orientation in an extinct agnathan. The presence of
fibrous extrinsic eye muscles confirms the degree of encephalization o
f the conodonts and is consistent with a cladistic position crownwards
of the myxinoids. The soft tissues are uniquely preserved as illite a
nd mixed-layer clay minerals.