The interstitial system of sandy beaches is lacunar and has its dimens
ions defined by the sand granulometry. It can be described by features
such as pore size, porosity, permeability, and water content. The mos
t important process occurring in this system, water filtration, is dri
ven by inputs of freshwater from groundwater discharge, and inputs of
seawater by tides, wave run-up, and subtidal wave pumping. Reflective
beaches have seawater input effected mainly by waves; they filter larg
e water volumes with short residence times. Dissipative beaches displa
y the opposite patterns, slowly filtering small volumes input by tides
. Flow patterns and their effects on interstitial climate are describe
d. The water table of the beach moves in response to groundwater disch
arge, tides, and waves and influences erosion/accretion processes on t
he beach face: a high water table promotes erosion. A series of moistu
re zones can be recognised from the dry surface sand at upper tide lev
els, to permanently saturated sand below the low tide water table, nam
ely: a stratum of dry sand, a stratum of retention, a stratum of resur
gence, and stratum of saturation. Interstitial chemistry is briefly de
scribed in terms of salinity changes, organic loads, oxygen content, a
nd nutrient cycling. It is concluded that the interstitial environment
of sandy beaches spans a continuum between physically and chemically
controlled extremes: the former condition occurs on coarse sand reflec
tive beaches, which experience low organic inputs and high filtration
rates of large water volumes - resulting in powerful hydrodynamic forc
es; the latter occurs on dissipative beaches of fine sand, which are s
ubject to high organic inputs and low filtration volumes - resulting i
n stagnation and steep vertical chemical gradients. Many intermediate
situations occur and these are more favourable to interstitial life th
an either of the extremes.