Th. Sparks et al., LIFE STRATEGIES OF CARABUS-PROBLEMATICUS (COLEOPTERA, CARABIDAE) AT DIFFERENT ALTITUDES ON SNOWDON, NORTH WALES, Journal of zoology, 236, 1995, pp. 1-10
In a study to examine the effect of climate on life cycles, a total of
408 Carabus problematicus were caught in pitfall traps at four altitu
de sites between 660 m and 1055 m on Snowdon, north Wales. Under the a
ssumption that the mandible condition of these beetles was related to
the age of the individuals, an objective estimate, which was both sex-
and size-independent, was used to separate the captures into two age
categories. The results from this estimation process suggested that be
etles from the different altitudes followed different life strategies;
low altitude beetles being predominantly annual and high altitude bee
tles predominantly biennial.