Ecological parapatry, in which pairs of largely allopatric taxa abut a
long common boundaries without hybridization, is often reported but se
ldom explained. A computer simulation model is developed that shows th
at parapatry between two species can be maintained by interspecific in
teraction on a dine of reducing ecological suitability for the competi
tively stronger species. In the model, a homogeneous environment requi
res much greater interaction strength to sustain parapatry than does a
heterogeneous environment with alternate regions of favorable and poo
r habitat. The heterogeneous environment of the model is intended to m
imic the environment near a well-studied parapatric boundary between t
wo reptile tick species.