Simulation of dry matter production by the explanatory glasshouse crop
growth model SUKAM (Gijzen, 1992, Simulation Monographs), based on SU
CROS87 (Spitters, Van Keulen and Van Kraalingen, 1989, Simulation and
systems management in crop protection), was validated for tomato. In t
he model, assimilation rates are calculated separately for shaded and
sunlit leaf area at different cumulative leaf area in the canopy, laki
ng into account the different interception of direct and diffuse compo
nents of light. Daily crop gross assimilation rate (P-gd) is computed
by integration of these rates over total crop leaf area and over the d
ay. Leaf photochemical efficiency and potential gross assimilation rat
e al saturating light depend on temperature and CO2 concentration and
are approximated as being identical in the whole canopy. Crop growth r
esults from P-gd minus maintenance respiration rate (R(m); dependent o
n temperature and crop dry weight), multiplied by the conversion effic
iency (carbohydrates to structural dry matter; C-f). Growth experiment
s (periodic destructive harvests) with different planting dates and pl
ant densities and two datasets from commercially grown crops, were use
d for model validation. Hourly averages for global radiation outside t
he glasshouse, glasshouse temperature and CO2 concentration, together
with measured leaf area index, dry matter distribution (for calculatio
n of C-f) and organ dry weights (for calculation of R(m)) were the inp
uts to the model. Dry matter production (both level and dynamic behavi
our) was simulated reasonably well for most experiments, but final dry
matter production was under-estimated by about 27% for the commercial
ly grown crops. At low irradiance and with large crop dry weight, grow
th rate was under-estimated, probably as a result of over-estimation o
f R(m). This could almost completely explain the large under-estimatio
n for the commercially grown crops, which had large dry weight. Final
dry matter production was over-estimated by 7-11% if daily averages in
stead of hourly input of climatic data were used. It is concluded that
SUKAM is a reliable model for simulating dry matter production in a t
omato crop, except for those situations where R(m) has a large influen
ce on crop growth rate (low irradiance and large crop dry weight). An
improved estimate of R(m) would take into account the influence of met
abolic activity. A preliminary attempt to relate maintenance costs to
relative growth rate (a measure for metabolic activity), showed promis
ing results.