Biogeographical and biodiversity studies in lowland Amazonian rain for
ests typically refer to observed or postulated distribution barriers s
uch as past unfavorable climates, mountains, rivers, and river floodpl
ains that divide the uniform tierra firme (noninundated) forest. Prese
nt-day ecological heterogeneity within tierra firme has hardly been di
scussed in,this context, although edaphic differences are known to aff
ect species distribution patterns in both inundated areas and tierra f
irme. Quantification of landscape heterogeneity in Peruvian lowland Am
azonia (500,000 kilometers squared), based on field studies and satell
ite image analysis, shows that Peruvian Amazonia is considerably more
heterogeneous than previously reported. These observations have implic
ations for the research, management, and conservation of Amazonian bio
diversity.