General guidelines on the quantitative representation of heterogeneity
in ecology are still lacking. Without a precise definition and sound
quantification of heterogeneity, statements involving the concept will
continue to be confusing, and comparative studies and hypothesis test
ing will be ineffective. To solve this problem we propose an operation
al definition of ecological heterogeneity that facilitates quantificat
ion based on the data types of concern. We use two examples to demonst
rate how our definition and quantification of spatial heterogeneity ca
n be applied in practice.