The influence of daylength on final leaf number was determined in four
field-sown wheat cultivars with no vernalisation requirement Leaf and
primordium appearance was monitored in plots sown at two-month interv
als during an annual cycle. The timing of the response to daylength va
ried substantially among cultivars. In one, final commitment of the la
st leaf primordium occurred close to the time of initiation of that pr
imordium while, in another cultivar, commitment did not occur until te
rminal spikelet or later. In the other two cultivars, commitment occur
red midway through spikelet initiation. Late timing of response to day
length is a useful adaptation that allows cultivars with no vernalisat
ion response sown in autumn to set more leaves than when sown in sprin
g, and hence to delay flowering. Based on the analysis, a simple itera
tive procedure was developed that successfully predicted final leaf nu
mber, the dares of appearance of the flag ligule and anthesis for one
of the cultivars grown in an independent experiment.