Global change research examines a variety of effects that changing env
ironmental conditions may have on ecosystems. One aspect, a decline in
species richness (biodiversity) observable in most ecosystems, has be
en less well studied, partly due to the complexity and long time neces
sary for experimentation. Model ecosystems (micro- or mesocosms), howe
ver, provide a feasible means for testing the many hypothetical relati
onships proposed to exist between diversity and ecosystem behaviour th
at observation and theory support. This article examines these hypothe
ses and shows how mesocosm studies conducted in the Ecotron, a control
led environmental facility, provide insights into this comples issue.