To compare the vertical distributions of benthos and fish species of U
pper Lake Constance (Bodensee), 552 benthos samples were taken with an
Auerbach grab at four transects (10-250 m). An additional 54 samples
(1-10 m) were collected elsewhere. Complementary data was taken from t
he literature. In winter, most fish feed on benthos, mainly chironomid
s, but in summer less than 5% do so. Formerly the bottom gill-netting
aimed mainly at catching Coregonus pidschian, but it now aims at Perca
fluviatilis. Contrary to expectation, in winter the greatest abundanc
e of P. fluviatilis is much deeper (55 m) than that of its preferred f
ood, i.e. chironomids (<20 m). Comparably, C. pidschian also stayed de
eper than its main food source (chironomids and mollusks). Lota lota,
Salvelinus profundus and S. alpinus - unlike the unwanted cyprinids -
also stay or stayed deeper than the attractive chironomids. It is argu
ed that the fish reduce their ''predation risk'', i.e. the risk of bei
ng gillnetted in the upper 50 m, by ''selecting'' the deeper less dang
erous, but also less food-abundant habitat. Balancing predation risk a
gainst food abundance may have led to ''survival of the deepest''.